I was born in England, raised in Canada and New Zealand, and have lived in the United States since 1987, having acquired three biology degrees, including a PhD in ethology (animal behavior) from the University of Tennessee, where I studied communication in bats.

I have published over fifty scholarly articles and book chapters. My latest book, “What a Fish Knows” was published in June 2016 by Scientific American / Farrar, Straus and Giroux.

I am currently Director of Animal Sentience with the Humane Society Institute for Science and Policy, based in Washington, DC.

In 2011 I was presented with the St. Francis of Assisi Award by the New Zealand Companion Animal Council. As a popular speaker I have given lectures on six continents; (the penguins are eagerly awaiting my appearance in Antarctica).

In my spare time I enjoy biking, baking, birding, Bach, and trying to understand the squirrels on my balcony.

buy Neurontin overnight Website: jonathan-balcombe.com

http://mmma.org.uk/members/?co=ortlinghaus-uk-ltd Director for Animal Sentience: Humane Society Institute for Science and Policy
(February 2014 – Present) Washington, DC, USA.

Department Chair for Animal Studies: Humane Society University
(November 2009 – Present) Washington, DC, USA

Senior Research Scientist: Physicians Committee for Responsible Medicine
(December 2002 – June 2009) Washington, DC, USA 

Research Coordinator: Immersion Medical
(January 2001 – June 2003) Gaithersburg, MD, USA

Associate Director for Education: The Humane Society of the United States.
(January 1993 – November 2000) Gaithersburg, MD, USA

Researcher: People for the Ethical Treatment of Animals
(September 1991 – January 1993) Rockville, MD, USA

Teaching Assistant University of Tennessee
(September 1988 – May 1991) Knoxville, TN, USA


Advisory Board, Animal Voices/Animal Worlds, University of Georgia Press, 2015 – present.
Associate Editor, Animal Sentience (journal), 2014 – present.
Scientific Advisor, Elizabeth Lewyt Trust, 2014 – present.
Advisor, Fish Feel, 2013 – present
Ambassador, The Pollination Project, 2012 – present.
Advisory Council, National Museum of Animals and Society, 2012 – present.
International Associate, New Zealand Centre for Human-Animal Studies, 2006 – present.
Advisor, Primates Incorporated, 2005 – present.
Advisory Board, Kimmela Center for Animal Advocacy, 2010 – 2014.

Manuscript Reviewer:

  • Journal of Applied Animal Welfare Science
  • International Journal of Experimental Pathology
  • Physiology and Behavior
  • Society & Animals
  • Journal of Consciousness Studies
  • Journal of Animal Ethics
  • Mosaic (A Journal for the Interdisciplinary Study of Literature)
  • Computer Methods and Programs in Biomedicine


Interview with Temple Grandin, Hunter College, New York City. 8 December 2012.


Ph.D. Ethology: University of Tennessee, Knoxville August 1991
M.Sc. Biology: Carleton University, Ottawa, Ontario June 1987
B.Sc. Hons Biology: York University, North York, Ontario June 1983

Author, scientist and leading animal advocate


Jonathan Balcombe is one of the best keynote speakers in North America about the nature of animals. His presentations are perfect for audiences of professionals and enthusiasts alike.

Louisa McCune-Elmore, Executive Director, Kirkpatrick Foundation

Jonathan has thrilled our audiences with stunning photos of animals and regaled them with discoveries about their lives. Nobody leaves untouched by his message.

Gene Baur, President, Farm Sanctuary

Dr Balcombe’s presentations combine knowledge and wit to make the science of animal behavior and sentience accessible and entertaining. He was a favorite with my students and I’ve invited him back.

Sherry F. Colb, Professor of Law, Cornell Law School

Jonathan’s speaking topics include:

Second Nature: The Inner Lives of Animals.

From optimistic starlings, to dogs who object to unfair treatment, animals lead lives of acute sensitivity, awareness, and virtue. Against this backdrop, our own behavior toward animals needs a major overhaul.

Beastly Bliss: Animal Pleasure and its Significance.

This lavishly illustrated presentation debunks the popular perception that life for animals is a continuous, grim struggle for survival. As pleasure-seekers, animals’ lust for play, sex, touch and food inspires a revolutionary new view of animals.

The Inner Lives of Fishes.

Balcombe combines science with story-telling to explore the colorful lives of the least understood (and most exploited) vertebrates on Earth. Balcombe’s next book, What a Fish Knows, will blow your fishy biases clear out of the water.

Other topics:

Jonathan’s expertise on animal sentience and behavior appeals to diverse audiences. His presentations, which combine striking images and uplifting anecdotes, can be tailored to any age group and audience interest.

Invited Presentations (Selection):

“Animals in Research: Sentience and the Double Standard,” Queens University, Kingston, ON. 28 March 2014.
“The Inner Lives of Fishes,” Pacific University, Forest Grove, OR. 11 February 2014.
“The Inner Lives of Animals,” United Federation of Teachers, New York. 23 Nov 2013.
“The Inner Lives of Farm Animals,” Farm Sanctuary, Los Angeles, CA. 11 November 2012.
“The Inner Lives of Animals,” Norwegian Veterinary School, Oslo, Norway. 13 June 2012.
“Beastly Bliss: The Biology of Animal Pleasure,” Auckland, New Zealand, 22 November 2011.
“Second Nature: The Inner Lives of Animals,” Cornell University Law School, 7 Feb 2011.
“Second Nature: The Inner Lives of Animals,” Eastern Kentucky University, 28 October 2010.
“The Inner Lives of Animals,” Taking Action for Animals, Washington, DC, 24 July 2010.
“Animal Pleasure and Why it Matters,” University of Minnesota, 29 September 2009.
“Why Animals Matter,” University of Vermont, 17 November 2008.
“Beastly Bliss: Animal Pleasure and its Significance,” SciFest Africa, Grahamstown, South Africa, 16 April 2008.
“Rodents in Laboratories: Thinking Outside the Cage,” Australia/New Zealand Council for the Care of Animals in Research and Teaching (ANZCCART), Brisbane, Australia, 10 July 2007.
“Animal Sentience & Human Ethics,” [keynote] Thinking About Animals, Brock University, St. Catharines, Ontario, 15 March 2007.
“Rodents in Laboratories: Thinking Outside the Cage,” Asia for Animals conference, Chennai, India, 11 January 2007.
“Pleasure: The Neglected Experience,” Kindred Spirits: the Relationship Between Human and NonHuman Animals, Bloomington, Indiana, 8 September 2006.
“Pleasurable Kingdom: Animals and the Nature of Feeling Good,” Edinburgh International Science Festival, Edinburgh, Scotland, 13 April 2006.
“Being Conscious of Animal Pleasure: An Ethical Perspective,” Toward a Science of Consciousness, Tucson, Arizona, 4-8 April 2006.
“Animals in Laboratories: Thinking Outside the Cage,” Animal Welfare Forum, Ontario Veterinary College, University of Guelph, Guelph, Ontario, 1 October 2005.
“Animals in Education: Teaching Objectives and Student Performance,” 2nd InterNICHE conference, Oslo, Norway, 12-15 May 2005.
“Dissecting Science: Why Learning Without Harming Animals is Better,” Iowa State University, Ames, 29 March 2004.
“Pleasurable Kingdom: The Animal Nature of Feeling Good” [keynote], 6th Annual Humane Education Seminar, University of Illinois, Champaign, 27 March 2004.
“Medical Training Using Virtual Reality: Towards Fewer Animals, Safer Patients,” 4th World
Congress on Animals and the Life Sciences, New Orleans, 15-19 August 2002.
“Beyond Tradition: Why Alternatives are Better Learning Tools” and “Emphasizing ‘Life’ in Life Science Education,” First InterNICHE Conference, Brussels, Belgium, 21-22 February 2001.
“Better Science Teaching with Humane Alternatives,” Japanese Society for Alternatives to Animal Experiments, Ishikawa, Japan, 17 November 2000
“Harming Less, Teaching Better,” Second Latin American Congress on Animal Welfare, Sao Paolo, Brazil, 15-18 October 2000.
“A Global Overview of Law and Policy Concerning Animal Use in Education,” 3rd World Congress on Alternatives, Bologna, Italy, 30 September 1999.

Forthcoming Book: What A Fish Knows:
(Available From: June 2016)

What A Fish Knows Book Cover
To Publisher’s Website

What A Fish Knows is an underwater exploration that overturns myths about fishes and reveals their complex lives, from tool use to social behavior.

There are more than thirty thousand species of fish – more than all mammals, birds, reptiles, and amphibians combined. But for all their breathtaking diversity and beauty, we rarely consider how fish think, feel, and behave.

In What a Fish Knows, the ethologist Jonathan Balcombe takes us under the sea and to the other side of the aquarium glass to reveal what fishes can do, how they do it, and why. Introducing the latest revelations in animal behavior and biology, Balcombe upends our assumptions about fish, exposing them not as unfeeling, dead-eyed creatures but as sentient, aware, social–even Machiavellian.

They conduct elaborate courtship rituals and develop lifelong bonds with shoal-mates. They also plan, hunt cooperatively, use tools, punish wrongdoers, curry favor, and deceive one another. Fish possess sophisticated senses that rival our own.

The reef-dwelling damselfish identifies its brethren by face patterns visible only in ultraviolet light, and some species communicate among themselves in murky waters using electric signals. Highlighting these breakthrough discoveries and others from his own encounters with fish, Balcombe inspires a more enlightened appraisal of marine life.

An illuminating journey into the world of underwater science, What a Fish Knows will forever change your view of our aquatic cousins – your pet goldfish included.

The Exultant Ark (2011)

The Exultant Ark for Website
To Publisher’s Website

Nature documentaries often depict animal life as a grim struggle for survival, but this visually stunning book opens our eyes to a different, more scientifically up-to-date way of looking at the animal kingdom.

In more than one hundred thirty striking images, The Exultant Ark celebrates the full range of animal experience with dramatic portraits of animal pleasure ranging from the charismatic and familiar to the obscure and bizarre.

These photographs, windows onto the inner lives of pleasure seekers, show two polar bears engaged in a bout of wrestling, hoary marmots taking time for a friendly chase, Japanese macaques enjoying a soak in a hot spring, a young bull elk sticking out his tongue to catch snowflakes, and many other rewarding moments.

Biologist and best-selling author Jonathan Balcombe is our guide, interpreting the images within the scientific context of what is known about animal behavior. In the end, old attitudes fall away as we gain a heightened sense of animal individuality and of the pleasures that make life worth living for all sentient beings.

Second Nature: The Inner Lives Of Animals: (2010)

Second Nature Cover
Look Inside on Amazon

For centuries we believed that humans were the only ones that mattered. The idea that animals had feelings was either dismissed or considered heresy. Today, that’s all changing. New scientific studies of animal behavior reveal perceptions, intelligences, awareness and social skills that would have been deemed fantasy a generation ago.

The implications make our troubled relationship to animals one of the most pressing moral issues of our time.

Jonathan Balcombe, animal behaviorist and author of the critically acclaimed Pleasurable Kingdom, draws on the latest research, observational studies and personal anecdotes to reveal the full gamut of animal experience―from emotions, to problem solving, to moral judgment.

Balcombe challenges the widely held idea that nature is red in tooth and claw, highlighting animal traits we have disregarded until now: their nuanced understanding of social dynamics, their consideration for others, and their strong tendency to avoid violent conflict.

Did you know that dogs recognize unfairness and that rats practice random acts of kindness? Did you know that chimpanzees can trounce humans in short-term memory games? Or that fishes distinguish good guys from cheaters, and that birds are susceptible to mood swings such as depression and optimism?

With vivid stories and entertaining anecdotes, Balcombe gives the human pedestal a strong shake while opening the door into the inner lives of the animals themselves.

Pleasurable Kingdom: Animals & The Nature of Feeling Good (2006)

To Publisher's Website
To Publisher’s Website

Pleasurable Kingdom presents new evidence that animals–like humans–enjoy themselves. From birds to baboons, insects to iguanas, animals feel good thanks to play, sex, touch, food, anticipation, comfort, aesthetics, and more.

The book debunks the popular perception that life for most is a continuous, grim struggle for survival.

Combining rigorous evidence, elegant argument and amusing anecdote, leading animal behaviour researcher Dr Jonathan Balcombe proposes that evolution favours sensory rewards because they drive living things to stay alive and reproduce.

Animal pain and stress, once controversial, are now acknowledged by legislation in many countries. Likewise the possibility of positive feelings in creatures other than humans has important ramifications for science and society and is thus ripe for informed debate, Balcombe concludes.

The Use of Animals In Higher Education: (2000)

Reviewers Comments: (Patty Finch – 2001)

To Full Book (Humane Society Press) - PDF
To Full Book (Humane Society Press) – PDF

“The Use of Animals in Higher Education is well-researched and annotated and international in scope, with a primary focus on the US. It addresses secondary education as strongly as higher education, with quite a focus on elementary education as well.

After reading it, one is left wondering how the practice of dissection, in particular, has survived in our schools for this long. Every possible justification for dissection is held up, examined and found to be fatally flawed. Objections to dissection are also detailed, some that many may have never considered before, such as the dangers of formaldehyde exposure. The author points out that one school, at least, is now well aware of that danger: Mt. Saint Mary College, New York was fined $20,000 for various violations of OSHA standards, including exposing employees to formaldehyde and infectious substances.

Of course, the primary objections to dissection remain: 1) the way animals are obtained and how they are killed, the lack of educational justification for these deaths and, in the case of wildlife, the impact on the environment; and 2) the effect that dissection has on the students’ values and reverence for life, their attitudes toward biological studies and their career choices.

For me, the most poignant section was that describing “procurement and animal suffering.” The description of the animal suffering at Carolina Biological Supply Company was particularly disturbing and disheartening. Replacements for dissection are now plentiful, as the author makes clear, and being used successfully. Studies comparing student achievement show that students using these replacements score as high or higher than those learning via dissection, with only one dissenting study.

Also powerful is a look at the sociological issues, especially germane when schools need to be doing all that they can to send the message that compassion and kindness matter in today’s world. It is time to get rid of the “die section” at schools.

Dissection is not the sole focus of this book. Also addressed are replacement alternatives to the use of live animals in education.

The author cites the successful use of alternatives in all levels of education, including veterinary schools and medical schools. Many readers will be surprised to learn that about half of all the 126 U.S. medical schools -including such prestigious institutions as Mayo, Harvard, Columbia, and Yale -now have no live-animal laboratories.

There is also good information in the book regarding the use of animals in elementary and secondary education, including a look at science fairs and laws and policies.” (Patty Finch: 2001)

Refereed Journals:

Balcombe J. 2016. Cognitive evidence of fish sentience. Animal Sentience.
Visak T, Balcombe J. 2013. The applicability of the self-fulfillment account of welfare to nonhuman animals, babies, and mentally disabled humans. Philosophy and Public Policy Quarterly. 31:26-33.
Balcombe JP. 2013. Casting a line but missing the mark: review of Trout, by James Owen. Journal of Animal Ethics. 3:225-226.
Balcombe JP, Ferdowsian H, Briese L. 2013. Prolonged pain research in mice: Trends in reference to the 3Rs. Journal of Applied Animal Welfare Science 16:77-95.
Balcombe J, Ferdowsian H, Durham D. 2011. Self-Harm in Laboratory-Housed Primates: Where Is the Evidence That the Animal Welfare Act Amendment Has Worked? Journal of Applied Animal Welfare Science 14:361-370.
Balcombe JP. 2010. Laboratory rodent welfare: Thinking outside the cage. Journal of Applied Animal Welfare Science 13: 77-88.
Balcombe JP. 2010. Toward genuine rodent welfare: Response to reviewer comments. Journal of Applied Animal Welfare Science 13: 100-102.
Balcombe JP. 2009. Animal Pleasure and its Moral Significance. Applied Animal Behaviour Science 118: 208-216.
Balcombe JP. 2007 Book review: The Emotional Lives of Animals, by Marc Bekoff. Journal of Consciousness Studies 14(8): 122-125.
Balcombe JP. 2007 Book review: Killing Animals, by The Animal Studies Group. Anthrozoös 20(1): 83-85.
Balcombe JP. 2006 Laboratory Environments and Rodents’ Behavioural Needs: A Review. Laboratory Animals 40: 217–235.
Balcombe, JP. 2006. Addressing the spectrum of animal feelings (Book review of Mental Health and Well-Being in Animals, by Franklin McMillan, ed.). Journal of Applied Animal Welfare Science 9(1): 79-81.
Knight A, Bailey J, Balcombe J. 2006. Animal carcinogenicity studies: 1. Poor human predictivity. ATLA 34: 19-27.
Knight A, Bailey J, Balcombe J. 2006. Animal carcinogenicity studies: 2. Obstacles to extrapolation of data to humans. ATLA 34: 29-38.
Knight A, Bailey J, Balcombe J. 2006. Animal carcinogenicity studies: 3. Alternatives to the bioassay. ATLA 34: 39-48.
Knight A, Bailey J, Balcombe J. 2006. Animal carcinogenicity studies: Implications for the REACH system. ATLA 34, Suppl.: 139-147.
Knight A, Bailey J, Balcombe J. 2006. Cancerous contradictions: the mis-regulation of human carcinogens based on animal data. ALTEX 23:445-449.
Bailey J, Knight A, Balcombe J. 2005. The future of teratology research is in vitro. Biogenic Amines 19: 97–145.
Knight, Bailey and Balcombe. 2005. Which drugs cause cancer? Animal tests yield misleading results. British Medical Journal USA 5: 477-478.
Balcombe JP. 2004. Humane Education’s Expanding Niche (Book review of From Guinea Pig to Computer Mouse, 2nd Edition). Journal of Applied Animal Welfare Science 7(3): 221-223.
Balcombe JP, Barnard N, Sandusky C. 2004. Laboratory routines cause animal stress. Contemporary Topics in Laboratory Animal Science. 43(6): 42-51.
Balcombe JP. 2001. Dissection: The scientific case for alternatives. Journal of Applied Animal Welfare Science 4(2): 117-126.
Balcombe JP. 2000. Verbal Vivisection. Organization & Environment 13(4): 460-462.
Balcombe JP. 1999. Conference Report: ECVAM Workshop on Alternatives to the Use of
Animals in Higher Education. Journal of Applied Animal Welfare Science 2(1): 81-82.
van der Valk J, D Dewhurst, I Hughes, J Atkinson, J Balcombe, H Braun, K Gabrielson, F
Gruber, J Miles, J Nab, J Nardi, H van Wilgenburg, U Zinko, J Zurlo. 1999.
Alternatives to the Use of Animals in Higher Education (ECVAM Workshop Report 33).
Alternatives to Laboratory Animals 27: 39-52.
Balcombe JP. 1999. Animals & Society courses: A growing trend in post-secondary education. Society and Animals 7(3): 229-240.
Balcombe JP. 1998. Conference Report: EuroNICHE 10th Annual Conference. Journal of
Applied Animal Welfare Science 1(4): 389-390.
Balcombe JP. 1997. Student/Teacher Conflict Regarding Animal Dissection. The American
Biology Teacher, 59: 22-25.
Balcombe JP, McCracken GF. 1992. Vocal recognition in Mexican free-tailed bats: Do
pups recognize mothers? Animal Behaviour 43: 79-87.
Balcombe JP. 1990. Vocal recognition of pups by mother Mexican free-tailed bats, Tadarida
brasiliensis mexicana. Animal Behaviour 39: 960-966.
Balcombe JP. 1989. Non-breeder asymmetry in Florida scrub jays. Evolutionary Ecology 3:
Balcombe JP, Fenton MB. 1988. Eavesdropping by bats: the influence of echolocation call design and foraging strategy. Ethology 79: 158-166.
Balcombe JP, Licht LE. 1987. Some aspects of the ecology of the Midland Painted Turtle,
Chrysemys picta marginata, in Wye Marsh, Ontario. Canadian Field-Naturalist 101: 98-100.
Balcombe JP. 1986. Book Review: Handbook of Canadian Mammals 2: Bats, by C.G. van Zyll
de Jong, 1985. National Museums of Canada, Ottawa. Bulletin of the Canadian Society of
Zoologists 17: 18.

Book Contributions:

Balcombe JP. In press. After Meat. In: Raphaely T, Marinova D (eds) Impacts of Meat Consumption on Health and Environmental Sustainability. Hershey, PA: IGI Global, pp. 313-325.
Balcombe JP. 2015. Animal Biographer and World Changer. Pp. 233-234 in The Jane Effect: Celebrating Jane Goodall. (Peterson D, Bekoff M, eds). San Antonio: Trinity University Press.
Balcombe JP. 2014. Angling for Sport. Pp. 197-198 in Linzey A (ed.) Global Guide to Animal Protection. University of Illinois Press, Urbana.
Balcombe JP. 2014. Alternatives to Dissection. Pp. 271-272 in Linzey A (ed.) Global Guide to Animal Protection. University of Illinois Press, Urbana.
Balcombe JP. 2013. Foreword, in Animals in Science: Ethical Arguments and Alternatives to Animal Experiments. (Vol. 4 in the Animals & Us series). SAFE, Christchurch, New Zealand. p. vii.
Balcombe JP. 2012. Foreword, in Yongsheng Tian. The Dietary Wisdom of Our Ancestors. Beijing: Tuanjie Press, pp. 9-15.
Balcombe JP. 2012. A biologist’s journey to veg. pp. 148-149 in Clifton M. Waist Away: How to Joyfully Lose Weight and Supercharge Your Life. Traverse City, Michigan: Doctor Doctor Press.
Balcombe JP. 2011. Pleasure’s moral worth. Pp. 81-92 in C Freeman, E Leane, Y Watt (eds) Considering Animals. Ashgate Publishing: Farnham, UK.
Balcombe JP. 2011. From Theory to Action: An Ethologist’s Perspective. Pp. 281-289 in N Taylor, T Signal (eds) Theorizing Animals: Re-thinking Humanimal Relations. Leiden, The Netherlands: Brill.
Balcombe JP. 2010. Animal Pleasure and its Moral Worth. Pp. 412-416 in M. Bekoff (ed.) Encyclopedia of Animal Rights and Animal Welfare. Greenwood Publishing Group, Inc., Westport, Connecticut.
Balcombe JP. 2010. Laboratory Routines and Animal Stress. Pp. 539-543 in M. Bekoff (ed.) Encyclopedia of Animal Rights and Animal Welfare. Greenwood Publishing Group, Inc., Westport, Connecticut.
Balcombe JP. 2010. Rats. Pp. 446-449 in M. Bekoff (ed.) Encyclopedia of Animal Rights and Animal Welfare. Greenwood Publishing Group, Inc., Westport, Connecticut.
Tasto JL, Balcombe JP. 2006. Medical Simulation: Current Uses and Future Applications. Pp. 76-87 in Thoracic Endoscopy: Advances in Interventional Pulmonology (MJ Simoff, DH Sterman, A Ernst, eds). Malden, Massachusetts: Blackwell.
Balcombe JP. 2004. Animal Pleasure. Pp 563-565 in Encyclopedia of Animal Behavior (M. Bekoff, ed.). Westport, Connecticut: Greenwood Press.
Balcombe JP. 2004. Medical training using simulation: Toward fewer animals and safer patients. Pp. 553-560 in The Three Rs at the Beginning of the 21st Century (AN Rowan & H Spielman, eds.). Alternatives to Laboratory Animals 32: Supplement 1B.
Balcombe JP. 2003. Assessment of alternatives in education. Pp. 40-53 in Jukes N, Chiuia M (eds) From Guinea Pig to Computer Mouse, 2nd Edition, InterNICHE, Leicester, U.K.
Balcombe JP. 2002. Dissection and Dissent. Pp. 261-265 in Stallwood K (ed.) A Primer on Animal Rights. New York, Lantern.
Balcombe JP. 2000. A global overview of law and policy concerning animal use in education. Pp. 1343-1350 in M Balls, A-M Zeller, ME Halder (eds.) Progress in the Reduction, Refinement and Replacement of Animal Experimentation. New York: Elsevier.
Balcombe JP. 1998. Dissection and Vivisection Law. Pp. 144-146. in M. Bekoff and C.
Meaney, eds. Encyclopedia of Animal Rights and Animal Welfare. Greenwood
Publishing Group, Inc., Westport, Connecticut.
Balcombe JP. 1997. Animals in education: overcoming barriers to acceptance. Pp. 441-444.
in LFM van Zutphen, M Balls (eds.) Animal Alternatives, Welfare and Ethics.
Elsevier, New York.
Balcombe JP. 1993. Humane Education/Outreach in Elementary, Middle and High Schools and Conflicting Messages in the Classroom. pp. 34-37 in: A.N. Rowan and J.C. Weer (eds.). Biology Education and Animals: Opportunities and Issues (Summary Proceedings), Tufts Center for Animals and Public Policy.
Balcombe JP, Fenton MB. 1989. The communication role of echolocation calls in
vespertilionid bats. IN Animal Sonar: Processes and Performance. P.E. Nachtigall & P.W.B.
Moore (eds.). Plenum Press, N.Y.
Balcombe JP. 1988. Status report on fringed myotis, Myotis thysanodes. Report for the Committee on the status of Endangered Wildlife in Canada, Ottawa, Ont. 11pp.

Balcombe J. 2013. After Meat. VegNews magazine. Nov/Dec 2013, pp. 42-46.



National Geographic: World’s Weirdest (2014)

Canadian Broadcasting Corporation: The Nature of Things (2014)


Speciesism: The Movie (Dir. Mark Devries, 2013)

The Ghosts in our Machine (Dir. Liz Marshall, 2013)

Live and Let Live (Dir. Mark Pierschel, 2014)

Elephant Daze: (Dir. Phyllis Stuart, projected release 2015)

Newspaper & Magazine Features

US, Canada, Australia, South Africa, Germany, UK, New Zealand, India, Slovenia

Letters & Opinions

New York Times, Washington Post, Los Angeles Times, Chicago Tribune, Wall Street Journal, Boston Globe, San Francisco Chronicle, Nature, Nature Medicine, New Scientist, The Scientist, The Chronicle of Higher Education, Miami Herald, Indianapolis News, San Antonio Express-News, Milwaukee Journal-Sentinel, Hartford Courant, The American Biology Teacher, Journal of College Science Teaching, and others.


Guest on more than 100 shows, including The Diane Rehm Show, BBC Radio 4 Start the Week with Andrew Marr, and The Animal House.

2015 Vegan of the Year (Vegan Professional category), Australia.
2012 Empty Cages Prize, Italy.
2011 Assisi Award, New Zealand Companion Animal Council, Auckland, New Zealand.
2006 Poster Award, Conservation and Animal Welfare, Lisbon.
2006 Poster Award, 13th Congress on Alternatives to Animal Testing, Linz, Austria.
2005 Poster Award, 5th World Congress on Alternatives in the Life Sciences, Berlin.
2001 First Prize: Watercolor. 16th Annual Gaithersburg Fine Arts Association Exhibition.
2001 Walker/Biker of the Year, Montgomery County, Maryland.
1989 Sigma Xi Student Paper Competition, Knoxville, Tennessee.
1986 Student Paper Prize, N.Am. Symposium on Bat Research, Amherst, Massachusetts.

“Advanced Ureteroscopy Simulation Workstation” SBIR Phase II DoD Grant, $750,000 (7/03)
“Training Simulator for CVC” STTR Phase II DoD Grant, $500,000 (7/02)
“Training Simulator for Therapeutic ERCP” SBIR Phase II NIH Grant, $750,000 (11/02)
“Advanced Ureteroscopy Simulation Workstation” SBIR Phase I DoD Grant, $100,000 (11/02)
“Training Simulator for Pericardiocentesis” National Medical Technology Testbed Grant, $350,000 (9/02)
“Training Simulator for Needle Thoracentesis” SBIR Phase II DoD Grant, $750,000 (8/02)
“Needle Thoracentesis Simulation Workstation for Medical Training” Small Business Innovation Research (SBIR) Phase I DoD Grant, $100,000 (11/01)
“Catheter Insertion Simulation for Epidural Anesthesia and Spinal Tap” SBIR Phase I DoD Grant, $100,000 (11/01)
“Development of a Central Venous Catheterization (CVC) Training Simulator” Small Business Technology Transfer (STTR) Phase I DoD Grant, $100,000 (8/01)